PRINCETON, N.J. -- Billy Ragone passed for one touchdown and ran for another while the Pennsylvania defense forced four turnovers -- including three interceptions -- as the Quakers outdueled Princeton 28-21 Saturday in a battle of first-place Ivy League foes.
Penn (4-4, 4-1) is still tied with next Saturday's opponent, Harvard, for first atop the Ivy League.
Each of Ragone's scores catapulted the Quakers into the lead, including a game-winning, 3-yard TD run with 2:59 to play that snapped a 21-21 tie.
Ragone's other score, a 14-yard TD pass to Ryan O'Malley, put the Quakers ahead 14-13 in the second quarter.
The turnovers undermined an effective offensive outing for the Tigers (4-4, 3-2), who out-passed Penn 319-96.
Penn's Lyle Marsh ran for 104 yards on 19 carries.
Florida State coach Jimbo Fisher hopes quarterback Sean Maguire is ready to go full speed by the middle of June.
Graduate transfer quarterback Davis Webb has officially joined California, giving the Golden Bears an experienced option to replace No. 1 overall NFL draft pick Jared Goff.
First-year football coach Dino Babers, hired by former Syracuse AD Mark Coyle, is on the committee that will search for Coyle's replacement.
Hugh Freeze has some major holes to fill on offense with the departures of Laremy Tunsil and Laquon Treadwell, but there is Chady Kelly and good depth.
After transferring in from Montana State, Dakota Prukop has a checklist he's working from as he prepares to compete for the Oregon job.
The destination for Week 6 of the Big 12 Ultimate Road Trip is unanimous, as all three conference reporters would head to the Red River Rivalry.