PRINCETON, N.J. -- Billy Ragone passed for one touchdown and ran for another while the Pennsylvania defense forced four turnovers -- including three interceptions -- as the Quakers outdueled Princeton 28-21 Saturday in a battle of first-place Ivy League foes.
Penn (4-4, 4-1) is still tied with next Saturday's opponent, Harvard, for first atop the Ivy League.
Each of Ragone's scores catapulted the Quakers into the lead, including a game-winning, 3-yard TD run with 2:59 to play that snapped a 21-21 tie.
Ragone's other score, a 14-yard TD pass to Ryan O'Malley, put the Quakers ahead 14-13 in the second quarter.
The turnovers undermined an effective offensive outing for the Tigers (4-4, 3-2), who out-passed Penn 319-96.
Penn's Lyle Marsh ran for 104 yards on 19 carries.
Ohio State remained No. 1 in the AP Top 25 and tied a poll record Sunday despite lagging support from voters. Meanwhile, No. 2 TCU and No. 3 Baylor edged forward and No. 4 Michigan State slipped.
Texas cornerback Kris Boyd has apologized for retweeting a Texas A&M fan during halftime of Texas' 50-7 loss to TCU on Saturday.
The Big 12 on Sunday acknowledged an officiating mistake that incorrectly awarded Oklahoma State a first down Saturday and said the officials involved would be disciplined.
Georgia coach Mark Richt says he's sticking with Greyson Lambert, who was just 10-for-24 for 86 yards in a loss to Alabama, as the Bulldogs starting quarterback.
Will Grier threw four touchdown passes to four receivers -- all in the first half -- and No. 25 Florida downed third-ranked Mississippi 38-10 on Saturday night.
Leonard Fournette highlighted his SEC-record third straight 200-yard game with a 75-yard touchdown run, and No. 9 LSU overcame Eastern Michigan 44-22 on Saturday night.